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       #Post#: 68912--------------------------------------------------
       More wolves-- "little dog gene" existed in wolves
       before dog domestication
   DIR By: guest114
       Date: January 31, 2022, 7:45 pm
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  HTML https://www.sciencedaily.com/releases/2022/01/220127114340.htm
       The paper is here:
  HTML https://www.cell.com/current-biology/fulltext/S0960-9822(21)01723-1<br
       />
       First off, let me clarify some points that are confused in the
       sciencedaily article:
       "..when the team looked at the DNA of a 54,000-year-old Siberian
       wolf (Canis lupus campestris) they found that it, too, possessed
       the growth hormone mutation. "
       1) IGF-1 (insulin-like growth factor 1, aka somatomedin C)  is
       not Growth Hormone (GF)-- it is stimulated by growth hormone.
       For instance, in humans Laron syndrome causes low IGF-1 because
       of growth hormone receptor defects.
       There's no known IGF-1 mutation that is linked to dwarfism in
       humans, probably because low or no IGF-1 tends to cause severe
       low blood sugar.  Knock-out mice models out show neurological
       defects and prenatal lethality.
       So, a viable IGF-1 polymorphism is not likely to be a mutation
       affecting its function, but something in how it is turned on.
       This is why they got somewhere AFTER :  "Jocelyn Plassais
       (@JocelynPlassais), a postdoc in geneticist Elaine Ostrander's
       lab, suggested that they search for sequences around the gene
       that were positioned backwards and confirm if any were present
       in other canids and ancient DNA."
       They were looking for inverted sequences around genes
       indicating inactivated or mutated promoters. Sometimes bits of
       chromosomes can get switched around (chromosomal inversions)
       because of very similar sequences at both ends.  Same for genes
       with lots of similar promoter sequences on both ends.
       "With this approach, their team found a reverse form of the
       insulin-like growth factor 1 (IGF1) gene with variants that
       correlated to dog body size."  That's the biggie.  It held up
       through analysis.   The IGF-1 gene was identified in small dogs
       over a decade ago, but establishing whether this existed in
       wolves took time.  They identified this gene in Arabian wolves,
       which are quite small BUT also have dog admixture, so it was not
       clear if this was from dogs OR what.
       "Or what" seems to have been proved-- it well predates all
       dogs.  It's the big dog variant that is new.  The actual Cell
       Article explains.
       This fits my suspicion we originally tamed a smaller wolf
       variant to dogs in the first place; Japanese wolves (which are
       descended from ancient Siberian wolves) was very small indeed,
       if bigger than Arabian wolves.
       
  HTML https://en.wikipedia.org/wiki/Japanese_wolf
       indeed, most dogs
       have a small trace of Japanese wolf after a post-domestication
       admixture into Japanese dog breeds.
       NOW to wolves.
       The Steppenwolf (that's the subspecies listed-- the Caspian
       wolf) is smaller than the Eurasian wolf, being around 74-88 lbs
       on average vs 85-110 lbs.  So that's significantly smaller,  but
       not tiny.  I'm not clear if they were smaller 54,000 years ago
       or not.  This specimen could be related to the ancestral pool
       leading to Japanese wolves.
       This is a nice find, proving this ancient DNA research proves
       the gene existed in wolves well before dogs were domesticated
       and probably was widespread at low frequencies, and then
       collapsed and went rare as wolf populations plummeted after the
       Pleistocene.  It also gives one possible marker to look for in
       ancient wolf DNA too when looking for the ancestor of dogs.
       
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